He clearly seems to be reigning it in on most of those---but not uncomfortable in the appearances to a general audience sense, which is what I thought was indented by the original post.
And I also didn't equate "uncomfortable" in any sense with "out of his element."
But anyway, personally uncomfortable with circumstances of a gig---sure, I bet a lot of times, even when "in his element." And seeming less than his usual self to us hardcore fans as he attempts to meld with others? Sometimes, but to a more general audience? I'm not so sure.

The history of his management (not Road Management! Hooray Bruce, Jim, Ed, Pete, Ed!) is truly a thread (or several) unto itself/themselves....